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C
Clade- This is a monophyletic group. Cladistics- (1)
The Journal of the Willi Hennig Society. (2) The scientific discipline
of classifying organisms according to the doctrine of Willi Hennig's 1966
book. (3) The belief that the organisation of taxa into groups can only
be done by the identification of synapomorphies
for that group.
Cladogram- A dendrogram
depicting the hypothesised branching order of a number of sequences. Cladograms
do not give any indication of temporal change, unlike a phylogram
which does.
Clock- also known as
a molecular clock. One of the most important assumptions that is
often made using molecular data is the existence of a clock. In other
words, there is sometimes an assumption being made either implicitly or
explicitly that the rate of change of residues in molecules is approximately
constant. Making this assumption is fraught with dangers, but a great
many empirical studies have shown that (with some errors) the assumption
of a clock often holds true. If there are some good reference points
(say, fossils or significant geological events) it is sometimes possible
to use molecular data to estimate the time of events for which there is
no other alternative.
Clustal- The name
of a program for performing automated multiple sequence alignment.
The original program was written by Desmond
G. Higgins when he worked as a post-doc in Paul Sharpe's lab in Dublin,
Ireland. In its present incarnation, Clustal is available as a text-only
program called ClustalW or as a graphical
program, with pull-down menus, called ClustalX.
Codon- This is a group
of three adjacent nucleotides that encode an amino acid. The identity
of the encoded amino acid can vary for a particular codon, depending on
the genetic code that is in use by an organism.
Congruence- Agreement
between characters or trees.
Consistency Index (CI)-
This is a measure of how well an individual character fits on a phylogenetic
tree. It is calculated by dividing the minimum possible number of
steps by the observed number of steps. If the minimum number of steps
is the same as the observed number of steps, then the character will have
a CI of 1.0. If a character is not completely compatible with a tree,
then it will have a CI value less than 1.0. The CI value of a tree
is the average CI value over all of the characters. There are two
problems with this value. It is expected to decrease with increasing
numbers of taxa and it is not clear what value is acceptably high or low.
Constraint- This
term usually refers to the placement of a caveat on the search algorithm,
so that it never examines trees that are pre-specified by the user as being
unwanted or unlikely. Constraint analysis is often used to speed-up
the search through tree-space. If a search is constrained so that
it does not look at improbable relationships (and therefore does not evaluate
these topologies), then it might proceed much more quickly. On the
other hand, constraint analysis might proceed by only finding those trees
that are incompatible with the constraint tree (reverse constraint analysis).
The latter analysis is used in order to place Decay Indices on the internal
branches of a tree.
Convergence- An
evolutionary event which occurs in two independent sequences. If the sequences
possess two different character states for a particular character, but
change results in each of these characters independently acquiring an identical
character state, this is said to be a convervence. Convergences differ
from parallellisms because the ancestral sequences are not the same. In
parallellisms,
the ancestral sequences are the same.
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Any
Questions? email: James Mcinerney
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